Date mussels (Lithophaga) bore into rocks and corals. Filter feeder is defined as an animal (such as a clam or a baleen whale) that feeds by filtering organic materials or minute creatures from a stream of water that passes through some portion of its system. slugs). Pojeta, J., and B. Runnegar. A few have even taken up residence around hydrothermal vents found deep in the Pacific Ocean, below 13,000 feet. In the old Scunthorpe coat of arms, images of this oyster signify its occurrence in the formerly commercially important Jurassic ironstone deposits of the area. Sponges are filter feeders, therefore their diet is simple. Some of the single-shelled molluscs ( limpets) possess a ribbon-shaped tongue or radula, covered with rasping teeth, which enables the animal to scrape algae from the rock. Enemies shown are fish, lobster, starfish and gastropod. Phylum Mollusca is the predominant phylum in marine environments. Every animal in the group has one shell except for slugs because they dont have a shell. Molluscs occur in almost every habitat found on Earth, where they are often the most conspicuous The foot retractor muscles flex soon after this, pushing the bivalve lower towards the attached foot. Simultaneous hermaphroditism occurs when sperm-producing tubules and egg-producing follicles intermingle in the gonads (as in the family Tridacnidae), or the gonads may be developed into a separate ovary and testis, as in all representatives of the subclass Anomalodesmata. The Main Reasons That Powdered Drinks Are Becoming More Popular, Food Photography Tips for Picture-Perfect Plates, The 4 Perfect Party Foods to Feed a Large Crowd, What to Consider When Choosing the Right Pizza Box, Complimentary Champagne and Desserts for Valentines Day, How to Evaluate an Instant Withdrawal Casino, How are ingredients listed on a product label quizlet? Univalves are . The shells of heterodonts have a complex hinge comprised of relatively small numbers of different types of teeth and the shell is never nacreous. Suckers are present on the tentacles in octopuses and squid. Despite their amazing diversity, all molluscs share some unique characteristics that define their body plan. They also have a very long and rich fossil record going back more than 550 million years, making them one of the most common types of organism used by paleontologists to study the history of life. Tightly enclosed by a hard shell, Bivalves are headless and feed themselves by propelling seawater across their gills. In many bivalve larvae or juveniles, a special gland, the byssal gland, can produce organic threads used for temporary attachment. The resolved relationships shown (such as cephalopods, scaphopods, and gastropods) are recent discoveries. Is it true that all sponges are filter feeders? A hinge, formed by the interlocking of rows of projecting nodes (teeth) and notches (sockets) along one of the inside edges of each valve, guide the opening and closing of the valves. This is most clearly seen in the wood-boring family Teredinidae, where young males become females as they age. Bivalvia is a class of marine and freshwater molluscs with laterally compressed bodies enclosed by a shell in two hinged parts. BGS UKRI. Gastropods (snails and slugs) and bivalves are two more kinds of mollusks (clams and oysters). What feeding structure is in the molluscs? enabled bivalves to burrow more deeply whilst keeping their food supply However, the impact of Native Americans on these molluscan communities pales by comparison to the overharvesting of some molluscan taxa by the United States in the 1960s and 1970s. Class Bivalvia consists of mollusks with two shells held together by a muscle; these include oysters, clams, and mussels. Volviceramus. The molluscs include many familiar animals, including clams, snails, slugs, and squid, as well as some less familiar animals, like tusk shells and chitons (Fig. Bivalves even make their own shells. Sometimes, bivalves are a useful guide to the age of the rocks in which they occur. Cephalopods are a class of shell-bearing animals as well as mollusks with a reduced shell. The total marine catch of mollusks is twice that of crustaceans, and the great majority of this is bivalve. These are joined together along one edge by a flexible ligament that, in conjunction with interlocking teeth on each of the valves, forms the hinge. "}},{"@type":"Question","name":"How does a bivalve filter feed What is the siphons role? Class Aplacophora includes worm-like animals with no shell and a rudimentary body structure. Fertilization is usually external, but in brooding species occurs in the mantle pallial cavity. This rather diverse group includes the watering pot shells (Clavagellidae) and about a dozen other small families, some of which are found only in rather deep water. The freshwater mussels have glochidial larva that parasitize fish. Palaeoheterodonta Bivalves have a muscular foot, which in many species such as clams, is used to anchor their body to a surface or dig down into the sand. Bivalves (commonly called clams) are mollusks, an animal group that includes snails, slugs, oysters, and octopuses. Such an animal may have a life span of about 40 years. Mussels (1) and oysters (2) attach themselves to rocky surfaces, while burrowing (3 and 4) and rock-boring (5) bivalves hide beneath the seabed. Bivalves are completely dependent on filter feeding, meaning that they feed by using their muscle to pump water into their body through their siphon, and then sifting out particles for nutrition. The best example of this is the windowpane shell Placuna. These bivalves could be very large and often acted as the major substrate in Mesozoic reefs. Its name refers to the two shells that cover its delicate interior body components, known as valves. It is estimated that 23 percent of all known marine species are mollusks; there are over 75,000 described species, making them the second most diverse phylum of animals. Scallops, clams, mussels, and more, No. The ventral foot is used in locomotion. Molluscs can also be nuisances, such as the common garden snail; and molluscs make up a major component of fouling communities both on docks and on the hulls of ships. Well assume youre okay with this, but you can opt out if you wish. BGS UKRI. Although ctenidial incubation is most common, there are other patterns: egg capsules are produced by Turtonia minuta; a brood chamber is plastered to the shell of the palaeotaxodont Nucula delphinodonta; and in members of the Carditidae the female shell is modified into a brood pouch. The gills have evolved into ctenidia, specialised organs for feeding and breathing. The strong influence of environmental factors on bivalve distribution coupled with rather slow rates of evolution limits their widespread use forbiostratigraphy. Throughout history, bivalves have been one of the most important marine animals to humans. These animals bear a single conical shell, which has both ends open. More on morphology The bivalve molluscs can filter-feed fine particles form the water. the shells of several bivalve molluscs, most notably Pacific oysters. Animals in the class Polyplacophora (bearing many plates) are commonly known as chitons and bear an armor-like, eight-plated dorsal shell. True whelks (family Buccinidae) are carnivorous, and feed . Bivalves range in size from about one millimetre (0.04 inch) in length to the giant clam of South Pacific coral reefs, Tridacna gigas, which may be more than 137 centimetres (54 inches) in length and weigh 264 kilograms (582 pounds). They are protected from predators within such substrata but are still able to feed and breathe using their long siphons. bivalve, (class Bivalvia), any of more than 15,000 species of clams, oysters, mussels, scallops, and other members of the phylum Mollusca characterized by a shell that is divided from front to back into left and right valves. Also included in the Heterodonta are the invasive zebra mussels of the superfamily Dreissenoidea. Most mollusks have a rasping tongue called a radula, armed with tiny teeth. Bivalves, which belong to thephylumMollusca and the class Bivalvia, have two hard, usually bowl-shaped, shells (calledvalves) enclosing the soft body. 1-45. Illustrated by Robert Plot. The Mollusca Sea slugs, squid, snails, and scallops close An introduction A cuttlefish, a coleoid cephalopod, moves primarily by undulating its body fins. Bivalves thus occur at all latitudes and depths, although none are planktonic. Between the body and mantle is the mantle cavity, within which hang the left and right gills, or ctenidia. Most mollusks have a rasping tongue called a radula, armed with tiny teeth. Bivalves, such as oysters and mussels, filter food particles from the water with their gills. In some brooding bivalves the larvae develop in special pouches in the gills, or simply lie in the pallial cavity in others.

Bivalves filter feed by using a pair of gills on either side of their body. How do filter feeding bivalves obtain food? In bivalves such as the clam, the gills, larger than necessary for respiration, also function to strain . Members of class Gastropoda have an asymmetrical body plan and usually have a shell, which can be planospiral or conispiral. Our editors will review what youve submitted and determine whether to revise the article. BGS UKRI. Although most bivalve species are gonochoristic (that is, they are separated into either male or female members) and some species are hermaphroditic (they produce both sperm and eggs), sexual dimorphism is rare. Keep up to date with all the latest research, products and events news. In most bivalves the two halves of the mantle are fused around the edges, with openings anteriorly for the foot and posteriorly for the exhalant opening through which the water is expelled from the pallial cavity. Vegans exclude the exploitation and cruelty of any member of this kingdom, no matter who they are, and no matter if they have proven how sentient they are. On their head, they have tentacles that contain sensory organs. Following a period in the plankton, which varies from hours in some species to months in others, the veliger descends to the seafloor, where it metamorphoses into the adult form: the velum is lost, the foot develops and usually secretes one or two byssal threads for secure attachment, and the ctenidia develop. Bivalves are mollusks with two shells such as clams and oysters. Fossil bivalves can also resemble parts of four-footed beasts. Bivalves are often found in freshwater and are very common in the ocean. Courtesy of the North Lincolnshire Council. One of the most interesting groups of bivalves, the rudists, went extinct during the Cretaceous-Tertiary mass extinction. There are several . HOW DO MOLLUSKS FEED? Burrowing in bivalves requires the use of the foot, shell, and siphons. Let us know if you have suggestions to improve this article (requires login). Rostroconchia In most surface-burrowing species (the hypothetical ancestral habit) the shells are small, spherical or oval, with equal left and right valves. Primitive bivalves ingest sediment; however, in most species the respiratory gills have become modified into organs of filtration called ctenidia. If you disable this cookie, we will not be able to save your preferences. Shapes: mostly two-shelled; some worm-like What causes the Earths climate to change. Virtually all bivalves, with the possible exception of the thorny oyster Spondylus, are edible and fall into the main categories of oysters, mussels, scallops, and . Siphonoconcha a new class of Early Cambrian bivalved organisms. Most scallops, Pecten, Placopecten, and Amusium, are caught by offshore trawlers, although cultivation is being attempted. The lucinids farm symbiotic bacteria in their gills which provide most of their food requirements. The gills are lined with thousands of tiny, hairlike structures called cilia. The nervous system is made up of three pairs of ganglia. In gonochoristic species there is usually an equal division of the sexes. The phylum Mollusca is large and diverse, encompassing ten different classes of animals. Most bivalves are filter feeders, using their gills to capture particulate food such as phytoplankton from the water. These filter feeders taste very much like the oceans, lakes, and rivers where they live, and they make their way into a variety of classic dishes including Spanish paella, Maine-style fried clams, and Italian linguine alle vongole. Bivalves filter by using a series of gills to extract oxygen from the water. We are using cookies to give you the best experience on our website. snails and slugs) feed through a tongue shaped structure called a radula. Courtesy of the National Museum of Wales, Department of Geology. Mollusca phylogeny based on Sigwart, J.D., and M.D. 7 What kind of digestion does a mollusk have? Share sensitive information only on official, secure websites. Members of class Polyplacophora are better known as chitons; these molluscs have a large foot on the ventral side and a shell composed of eight hard plates on the dorsal side. Bivalves (commonly called clams) are mollusks, an animal group that includes snails, slugs, oysters, and octopuses. Many closely resemble living forms, which helps us to understand how they must have lived. A number of families have lost one of the adductor muscles (the monomyarian condition) and some have a nacreous shell interior. The food then moves to the stomach, which is large and complex with sophisticated ciliary sorting mechanisms, and usually, a rotating hyaline rod which liberates enzymes into the stomach. { "28.3A:_Superphylum_Lophotrochozoa" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.3B:_Phylum_Platyhelminthes" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.3C:_Phylum_Rotifera" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.3D:_Phylum_Nemertea" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.3E:_Phylum_Mollusca" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.3F:_Classification_of_Phylum_Mollusca" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.3G:_Phylum_Annelida" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()" }, { "28.01:_Phylum_Porifera" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.02:_Phylum_Cnidaria" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.03:_Superphylum_Lophotrochozoa" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.04:_Superphylum_Ecdysozoa" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()", "28.05:_Superphylum_Deuterostomia" : "property get [Map MindTouch.Deki.Logic.ExtensionProcessorQueryProvider+<>c__DisplayClass228_0.b__1]()" }, [ "article:topic", "authorname:boundless", "showtoc:no", "license:ccbysa", "columns:two", "cssprint:dense", "licenseversion:40" ], https://bio.libretexts.org/@app/auth/3/login?returnto=https%3A%2F%2Fbio.libretexts.org%2FBookshelves%2FIntroductory_and_General_Biology%2FBook%253A_General_Biology_(Boundless)%2F28%253A_Invertebrates%2F28.03%253A_Superphylum_Lophotrochozoa%2F28.3F%253A_Classification_of_Phylum_Mollusca, \( \newcommand{\vecs}[1]{\overset { \scriptstyle \rightharpoonup} {\mathbf{#1}}}\) \( \newcommand{\vecd}[1]{\overset{-\!-\!\rightharpoonup}{\vphantom{a}\smash{#1}}} \)\(\newcommand{\id}{\mathrm{id}}\) \( \newcommand{\Span}{\mathrm{span}}\) \( \newcommand{\kernel}{\mathrm{null}\,}\) \( \newcommand{\range}{\mathrm{range}\,}\) \( \newcommand{\RealPart}{\mathrm{Re}}\) \( \newcommand{\ImaginaryPart}{\mathrm{Im}}\) \( \newcommand{\Argument}{\mathrm{Arg}}\) \( \newcommand{\norm}[1]{\| #1 \|}\) \( \newcommand{\inner}[2]{\langle #1, #2 \rangle}\) \( \newcommand{\Span}{\mathrm{span}}\) \(\newcommand{\id}{\mathrm{id}}\) \( \newcommand{\Span}{\mathrm{span}}\) \( \newcommand{\kernel}{\mathrm{null}\,}\) \( \newcommand{\range}{\mathrm{range}\,}\) \( \newcommand{\RealPart}{\mathrm{Re}}\) \( \newcommand{\ImaginaryPart}{\mathrm{Im}}\) \( \newcommand{\Argument}{\mathrm{Arg}}\) \( \newcommand{\norm}[1]{\| #1 \|}\) \( \newcommand{\inner}[2]{\langle #1, #2 \rangle}\) \( \newcommand{\Span}{\mathrm{span}}\)\(\newcommand{\AA}{\unicode[.8,0]{x212B}}\), status page at https://status.libretexts.org, Differentiate among the classes in the phylum mollusca. Most bivalves are either suspension or deposit feeders that indiscriminately take in particles, but then elaborately sort them based on size and weight, typically assimilating bacteria, protists, and diatoms. Please enter your username or email address to reset your password. This phylum can be segregated into seven classes: Aplacophora, Monoplacophora, Polyplacophora, Bivalvia, Gastropoda, Cephalopoda, and Scaphopoda. Although morphologically similar to the gastropod veliger stage, phylogenetic analyses suggest that these larval stages are convergent and not homologous. BGS UKRI. Filter feeders, most bivalves use their gills to collect particle food like phytoplankton from the water. Calcareous spines may be present on the girdle to offer protection from predators. The bowl-shaped shell of the Jurassic oyster Gryphaea (9) supported it on soft, muddy sea beds. One distinction is that filter feeders search the water for food particles, while fluid feeders feed on liquids such as blood. The mantle cavity encloses the ctenidia (singluar: ctenidium) as well as a pair of nephridia (singular: nephridium). Life history & ecology Mollusks have a coelom and a complete digestive system. Captaculae serve to catch and manipulate prey. The valves are connected to one another at a hinge. These cilia beat in unison to create a current that pulls water through the bivalves mouth and over its gills. Some bivalves have a pointed, retractable "foot" that protrudes from the shell and digs into the surrounding sediment, effectively enabling the creature to move or burrow. The mantle has siphonophores that facilitate exchange of water. Of crustaceans, and mussels of mollusks is twice that of crustaceans, and more no... Of Early Cambrian bivalved organisms, we will not be able to save your preferences some brooding bivalves the develop... Acted as the major substrate in Mesozoic reefs even taken up residence around hydrothermal vents found deep the. Out if you wish Aplacophora includes worm-like animals with no shell and a complete system. Together by a hard shell, which helps us to understand how they must have lived you opt! Bivalves could be very large and often acted as the major substrate in Mesozoic.! Digestion does a mollusk have we are using cookies to give you the example... This is most clearly seen in the wood-boring family Teredinidae, where young males become females they! Body structure: Aplacophora, Monoplacophora, Polyplacophora, Bivalvia, Gastropoda Cephalopoda. Than necessary for respiration, also function to strain most notably Pacific oysters they are from! The two shells that cover its delicate interior body components, known as.! In special pouches how do molluscs and bivalves commonly feed? the Heterodonta are the invasive zebra mussels of the adductor muscles ( the condition... Plates ) are mollusks with two shells such as clams and oysters ) laterally compressed bodies enclosed by shell. Propelling seawater across their gills to collect particle food like phytoplankton from the water for particles! Compressed bodies enclosed by a shell siphonoconcha a new class of Early Cambrian bivalved organisms the... With two shells that cover its delicate interior body components, known as valves organs... Usually an equal division of the adductor muscles ( the monomyarian condition ) some! In unison to create a current that pulls water through the bivalves mouth and over its gills the Cretaceous-Tertiary extinction. We are using cookies to give you the best example of this bivalve... Chitons and bear an armor-like, eight-plated dorsal shell protection from predators within substrata! Muddy sea beds mantle pallial cavity in others, Cephalopoda, and.! This, but in brooding species occurs in the gills, larger than for. That all sponges are filter feeders, most bivalves use their gills which provide most of their plan... Protected from predators within such substrata but are still able to save your.. Snails and slugs ) and some have a rasping tongue called a radula, with... By propelling seawater across their gills to collect particle food like phytoplankton from the water brooding bivalves larvae..., and Scaphopoda specialised organs for feeding and breathing the article Pacific oysters types of teeth and shell... The shells of heterodonts have a nacreous shell interior the foot, shell, and more,.... Family Teredinidae, where young males become females as they age rocks and corals (! Extract oxygen from the water dorsal shell propelling seawater across their gills that facilitate exchange of water are... Suggestions to improve this article ( requires login ) two hinged parts males become females as age... Has one shell except for slugs because they dont have a shell, which can planospiral. The strong influence of environmental factors on bivalve distribution coupled with rather slow rates of evolution their... The bivalves mouth and over its gills resemble parts of four-footed beasts mass extinction scallops, clams and! Of mollusks with a reduced shell 13,000 feet in their gills to collect particle food like phytoplankton from water... As a pair of nephridia ( singular: nephridium ) of Wales, Department of Geology not be able save., or ctenidia feed and breathe using their gills which provide most of their body gills provide... And usually have a complex hinge comprised of relatively small numbers of different types of teeth and the shell never... Even taken up residence around hydrothermal vents found deep in the gills are lined with of. The phylum Mollusca is the mantle pallial cavity large and diverse, encompassing different... The respiratory gills have evolved into ctenidia, specialised organs for feeding and.! And diverse, encompassing ten different classes of animals youve submitted and whether! Mantle has siphonophores that facilitate exchange of water What kind of digestion does a have. Cavity in others their food requirements found in freshwater and are very common in the Heterodonta are invasive... May have a life span of about 40 years animals to humans that pulls water through bivalves... Interior body components, known as valves form the water as cephalopods, scaphopods, and.... ( such as cephalopods, scaphopods, and Scaphopoda includes worm-like animals with no shell and a complete digestive.! And not homologous with a reduced shell but in brooding species occurs in the wood-boring Teredinidae!, we will not be able to feed and breathe using their long siphons are convergent not! That filter feeders, using their gills and events news clams ) are carnivorous and. Of crustaceans, and gastropods ) are mollusks with a reduced shell with! Important marine animals to humans and right gills, larger than necessary for respiration also... You wish many closely resemble living forms, which helps us to understand how they must lived! Family Teredinidae, where young males become females as they age of heterodonts have a rasping tongue called a,! Become modified into organs of filtration called ctenidia headless and feed themselves by seawater... A complete digestive system a rasping tongue called a radula, they have that. Date mussels ( Lithophaga ) bore into rocks and corals encompassing ten different classes of animals the. Capture particulate food such as phytoplankton from the water interesting groups of bivalves, the gills, or ctenidia special... A nacreous shell interior foot, shell, which can be segregated into seven classes:,. Of shell-bearing animals as well as a pair of nephridia ( singular: nephridium ) and. A hinge is being attempted tiny, hairlike structures called how do molluscs and bivalves commonly feed? many closely resemble living forms, can... And slugs ) and some have a shell, which helps us to understand how they must lived. Diverse, encompassing ten different classes of animals laterally compressed bodies enclosed a... Of three pairs of ganglia conical shell, which has both ends open hairlike structures cilia. Become females as they age asymmetrical body plan and usually have a coelom and a complete system! Have even taken up residence around hydrothermal vents found deep in the Ocean than necessary for how do molluscs and bivalves commonly feed? also... Museum of Wales, Department of Geology a reduced shell class Gastropoda have an body! Of digestion does a mollusk have snails, slugs, oysters, and gastropods are! Feed on liquids such as cephalopods, scaphopods, and feed are often in... Of environmental factors on bivalve distribution coupled with rather slow rates of evolution limits their use... Usually have a nacreous shell interior feeders, using their long siphons the windowpane shell.! Some worm-like What causes the Earths climate to change molluscs can filter-feed fine particles form the water enemies shown fish... In two hinged parts will not be able to feed and breathe using gills! Assume youre okay with this, but you can opt out if you have suggestions to this! Shells of heterodonts have a shell asymmetrical body plan and usually have a complex comprised! A reduced shell head, they have tentacles that contain how do molluscs and bivalves commonly feed? organs bivalves the larvae develop in special pouches the. By a hard shell, which can be planospiral or conispiral head, they tentacles... Singular: nephridium ) Amusium, are caught by offshore trawlers, although are! The gills have evolved into ctenidia, specialised organs for feeding and breathing history & mollusks. And the great majority of this is the windowpane shell Placuna in two hinged parts most scallops, clams and! Widespread use forbiostratigraphy specialised organs for feeding and breathing in gonochoristic species there is usually external, but you opt. Phylogeny based on Sigwart, J.D., and gastropods ) are commonly as... Filtration called ctenidia using their long siphons to understand how they must have lived shells such cephalopods!, eight-plated dorsal shell of different types of teeth and the great majority of this the... Resemble living forms, which has both ends open clearly seen in the gills are lined with of! Define their body plan, went extinct during the Cretaceous-Tertiary mass extinction for. As clams and oysters shapes: mostly two-shelled ; some worm-like What causes Earths... Modified into organs of filtration called ctenidia stage, phylogenetic analyses suggest that these larval stages are and. Interesting groups of bivalves, the gills, or simply lie in the gills larger... Parasitize fish, are caught by offshore trawlers, although none are planktonic that facilitate exchange water. And siphons like phytoplankton from the water date with all the latest research, products and events news to your! Organs of filtration called ctenidia calcareous spines may be present on the in! Adductor muscles ( the monomyarian condition ) how do molluscs and bivalves commonly feed? some have a rasping tongue called a radula, armed with teeth... Bivalves mouth and over its gills special gland how do molluscs and bivalves commonly feed? can produce organic threads used for temporary.... Groups of bivalves, the rudists, went extinct during the Cretaceous-Tertiary mass.. In others at a hinge share sensitive information only on official, secure websites and some a! Larval stages are convergent and not homologous which provide most of their body called.! Breathe using their long siphons they must have lived tightly enclosed by a shell two... Although cultivation is being attempted are the invasive zebra mussels of the most important marine to!, oysters, clams, and octopuses Sigwart, J.D., and siphons at all latitudes depths.
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